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Temporal range: Jurassic or Cretaceous[1]
Scientific classification edit
Kingdom: Plantae
Clade: Tracheophytes
Division: Polypodiophyta
Class: Polypodiopsida
Order: Salviniales
Family: Marsileaceae
Genus: Regnellites
T.Yamada & M.Kato
R. nagashimae
Binomial name
Regnellites nagashimae
T.Yamada & M.Kato

Regnellites nagashimae is the oldest known fossil belonging to the fern family Marsileaceae. It comes from rocks of the Upper Jurassic or Lower Cretaceous Kiyosu-e Formation of Japan.[1] The fossils include leaves with visible veins, as well as sporocarps.


  1. ^ a b Yamada, Toshihiro; Masahiro Kato (2002). "Regnellites nagashimae gen. et sp. nov., the Oldest Macrofossil of Marsileaceae, from the Upper Jurassic to Lower Cretaceous of Western Japan". International Journal of Plant Sciences. 163 (5): 715–723. doi:10.1086/342036. S2CID 84631686.

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