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A large orthalicid from eastern Peru (probably of genus Sultana), recovering from substantial shell damage.
Scientific classification e
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
Subclass: Heterobranchia
Superorder: Eupulmonata
Order: Stylommatophora
Informal group: Sigmurethra
Superfamily: Orthalicoidea
Albers, 1860

See text

~1750 taxon names are available

The Orthalicoidea are a superfamily of air-breathing land snails, terrestrial gastropod mollusks in the informal group Sigmurethra.[3]


The Orthalicoidea is a dominant faunal element in the Neotropics, but also has a number of genera with a Gondwanan distribution.[1]


2005 taxonomy

This taxonomy, as accepted by Bouchet & Rocroi, was based on the study by Nordsieck, published in 1986,[4] and the classification as formulated by Schileyko in 1999.[5] However Bouchet & Rocroi diverged from this classification in uniting the families Bulimulidae and Orthalacidae. They also considered the family Placostylidae as a distinct family from Orthalicidae, and the family Coelociontidae a distinct family from Urocoptidae. However, the position of the families Megaspiridae and Grangerellidae is doubtful.[3]

The following seven families have been recognized in the taxonomy of Bouchet & Rocroi (2005):

2008 taxonomy

Uit de Weerd moved two families Urocoptidae and Cerionidae to newly established superfamily Urocoptoidea based on molecular phylogeny research in 2008.[6]

2009 taxonomy

Family Prestonellidae van Bruggen, 1978 - A study by Herbert and Mitchell, published in 2009, places this enigmatic family, of which the phylogenetic relationships were previously unknown, in the Gondwanan superfamily Orthalicoidea,[7] while it was previously tentatively placed as a synonym of Aillyidae.[3]

2010 taxonomy

Breure et al. (2010)[8] moved Prestonella and Bothriembryon to Placostylidae and they elevated Bulimulinae to Bulimulidae, Odontostomini to Odontostomidae, Amphibuliminae to Amphibulimidae.[8] They also removed Coelociontidae from Orthalicoidea.[8]

2012 taxonomy

Breure & Romero (2012)[9] confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae, renamed Placostylidae to Bothriembryontidae.[9] Therefore, there are recognized seven extant[9] families and one extinct[3] family within Orthalicoidea:


This article incorporates CC-BY-3.0 text from the reference[1]

  1. ^ a b c Breure A. S. H. (2011). "Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with description of two new species". ZooKeys 101: 1-50. doi:10.3897/zookeys.101.1133.
  2. ^ Cite error: The named reference WoRMS was invoked but never defined (see the help page).
  3. ^ a b c d Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
  4. ^ Nordsieck H. (1986). "The system of the Stylommatophora (Gastropoda), with special regard to the systematic position of the Clausiliidae, II. Importance of the shell and distribution". Archiv für Molluskenkunde. 117 (1–3): 93–116.
  5. ^ Schileyko A. A. (1998–2003). "Treatise on recent terrestrial pulmonate molluscs". Ruthenica. suppl. 2: 1–1626.
  6. ^ Uit de Weerd D. R. (2008). "Delimitation and phylogenetics of the highly diverse land snail family Urocoptidae (Gastropoda, Pulmonata) based on 28S rRNA sequence data: A reunion with Cerion". Journal of Molluscan Studies 74: 317-329. doi:10.1093/mollus/eyn023.
  7. ^ Herbert D. G. & Mitchell A. (2009). "Phylogenetic relationships of the enigmatic land snail genus Prestonella: the missing African element in the Gondwanan superfamily Orthalicoidea (Mollusca: Stylommatophora)". Zoological Journal of the Linnean Society. 96 (1): 203–221. doi:10.1111/j.1095-8312.2008.01109.x. Archived from the original on 2013-01-05.
  8. ^ a b c Breure A. S. H., Groenenberg D. S. J. & Schilthuizen M. (2010). "New insights in the phylogenetic relations within the Orthalicoidea (Gastropoda, Stylommatophora) based on 28S sequence data". Basteria 74(1-3): 25-31.
  9. ^ a b c Breure A. S. H. & Romero P. (2012). "Support and surprises: molecular phylogeny of the land snail superfamily Orthalicoidea using a three-locus gene analysis with a divergence time analysis and ancestral area reconstruction (Gastropoda: Stylommatophora)". Archiv für Molluskenkunde: International Journal of Malacology 141(1): 1-20. doi:10.1127/arch.moll/1869-0963/141/001-020.

Further reading

  • Borrero F. J. & Breure A. S. H. (Submitted). "The Orthalicoidea (Mollusca: Gastropoda: Stylommatophora) from Colombia and adjacent areas: a review of taxonomy and biogeography. 1. Dryptus Albers, 1860 and Plekocheilus Guilding, 1828".
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